The Mona Lisa smile has managed to bewitch us for 500 years. It is the culmination of a lifetime spent studying art, science and optics.
Leonardo's notebook shows his dissecting the human face to understand how every muscle and nerve touched the lips. He dissected the human eye on cadavers to understand that the retina's centre sees detail, but the edges see shadows and shape better.
If you look directly at the Mona Lisa's smile, the corners of the lips turn downward slightly, but shadows and light make it seem like it's turning upwards.
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