During typical running at a moderate effort, heart rate, oxygen uptake (which is a proxy for how much energy you’re burning), and perceived effort are all pretty similar on the treadmill and overground.
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Studies showed that running on a flat treadmill burned about 4 percent less energy, but that difference could be eliminated by setting the treadmill incline to 1 percent.
Air resistance depends on how fast you’re going. If you’re running really fast, air resistance takes a bigger...
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